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Some useful equations for field-sampling calculations:
population estimate = mean number of organisms per quadrat x (total area of habitat / area of one quadrat)
percentage cover = (number of squares in a quadrat containing the organism / total number of squares in the quadrat) x 100
percentage energy (or biomass) transfer between trophic levels = (energy available to the next trophic level / energy available at the current trophic level) x 100
In most ecosystems, only around 10% of the energy taken in by an organism is transferred to the trophic level above it; the rest is lost, mainly through respiration, movement, and materials that are not eaten or not digested.